Essential Parts of Flower: Androecium - Development of Anther

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Notes

Development of Anther:

Stages in the development of anther

(a) Anther primordium

(b) Differentiation of archesporial cell

(c) Formation of parietal and sporogenous cell

(d) Formation of wall layers

(e) Sporogenous stage

(f) Pollen tetrad stage

(g) Microspore stage

(h) Pollen grain stage

  • A very young anther develops as a homogenous mass of cells surrounded by an epidermis. 
  • During its development, the anther assumes a four-lobed structure. In each lobe, a row or a few rows of hypodermal cells becomes enlarged with conspicuous nuclei. This functions as archesporium.
  • The archesporial cells divide by periclinal divisions to form primary parietal cells towards the epidermis and primary sporogenous cells towards the inner side of the anther. 
  • The primary parietal cells undergo a series of periclinal and anticlinal division and form 2-5 layers of anther walls composed of endothecium, middle layers and tapetum, from periphery to centre whereas the sporogenous cell undergoes fewer divisions and forms the microspores, microsporocytes or pollen mother cells (PMC).
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